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Penkman, K. E. H., R. C. Preece, D. R. Bridgland, D. H. Keen, T. Meijer, S. A. Parfitt, T. S. White, and M. J. Collins. 2011. A chronological framework for the British Quaternary based on Bithynia opercula. Nature, 476: 446-449.
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